Wisteria Lane is about to get a lot more crowded.

Dana Delany (Kidnapped), Nathan Fillion (Drive), and Lyndsy Fonseca (Heroes) will join the cast of Desperate Housewives for the series' fourth season. Delany and Fillion will play a woman and her significantly younger husband who move to Wisteria Lane with the woman's daughter (Fonseca).

The move is an interesting one, not just for the casting of Fillion (who might make me reconsider giving up this once great series... for a second), but for that of Delany. Delany was originally the forerunner for the role of Bree (Marcia Cross), but turned down the pilot instead. Series creator Marc Cherry is said to be the one who courted Delany in the first place and is behind the drive to get her to finally sign on to Desperate Housewives.

If Delany brings some of the vicious energy of her role in Pasadena to the table, the ladies of Wisteria Lane won't know what hit them.

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In other casting news, Stephanie Niznik (Everwood) has joined the cast of the CW's family drama Life is Wild, which follows a veterinarian (Hidden Palms' D.W. Moffett, who takes over for Brett Cullen) who packs up his extended brood of children to leave behind Manhattan for a South African game reserve.

Niznik, who played Nina Feeney on Everwood, will star as the vet's second wife, a corporate lawyer. She replaces Judith Hoag, who originated the role in the pilot.

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Jace is an LA-based television development and acquisitions exec who watches way too much television for his own good and would love a TiVo for every room in the house. (He’s halfway there.) His blog, Televisionary, can be found at televisionaryblog.com.

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